ResearchIn-Press PreviewImmunologyPulmonology
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10.1172/jci.insight.202442
Aberrant mucin expression and keratinization distinguishing severe from mild asthma revealed by interpretable machine learning
Sagar L. Kale,1 Augusta M. Vincent,2 Mark A. Ross,3 Isha Mehta,2 Michael J. Calderon,3 Richard P. Ramonell,1 Himanshu Setya,4 Jessica C. McCreary-Partyka,1 Huijuan Yuan,1 Stephanie A. Christenson,5 Prescott G. Woodruff,5 Mario Castro,6 Kaharu Sumino,7 Nizar N. Jarjour,8 Loren C. Denlinger,8 Benjamin Gaston,9 Eugene R. Bleecker,10 Deborah A. Meyers,10 Wendy C. Moore,11 Elliot Israel,12 Bruce D. Levy,12 David Mauger,13 Serpil Erzurum,14 Anthony Newbrough,1 Taylor J. Nee,1 Prabir Ray,1 Claudette M. St. Croix,3 Sally E. Wenzel,1 Jishnu Das,15 Anuradha Ray,1 and Marc C. Gauthier1
1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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1Division of Pulmonary, Allergy, Sleep, and Critical Care Medicine, Departme, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
2Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, United States of America
3Department of Cell Biology, University of Pittsburgh, Pittsburgh, United States of America
4University of Pittsburgh School of Medicine, Pittsburgh, United States of America
5Division of Pulmonary, Critical Care, Sleep and Allergy, Department of Medi, UCSF, San Francisco, United States of America
6Division of Pulmonary, Critical Care and Sleep Medicine, University of Kansas School of Medicine, Kansas City, United States of America
7Division of Pulmonary and Critical Care Medicine, Washington University School of Medicine, St. Louis, United States of America
8Division of Allergy, Pulmonary and Critical Care Medicine, University of Wisconsin School of Medicine, Madison, United States of America
9Riley Hospital for Children and Indiana University School of Medicine, Indianapolis, United States of America
10Division of Genetics, Genomics and Precision Medicine, Department of Medici, University of Arizona, Tucson, United States of America
11Section on Pulmonary, Critical Care, Allergy & Immunologic Diseases, Wake Forest School of Medicine, Winston-Salem, United States of America
12Pulmonary and Critical Care Medicine Division, Department of Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, United States of America
13Division of Statistics and Bioinformatics, Department of Public Health Scie, Pennsylvania State University, University Park, United States of America
14Lerner Research Institute, Respiratory Institute, Cleveland Clinic, Cleveland, United States of America
15Center for Systems Immunology, University of Pittsburgh, Pittsburgh, United States of America
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Published July 30, 2026 - More info
Copyright © 2026, Kale et al. This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.
